Current Lab Members   |   Lab Alumni



Ben Ewen-Campen

Graduate Student

Cassandra Extavour

Ben graduated from Swarthmore College in 2006, where he studied Biology and English. Although he was initially drawn to bird biology in Alaska, he soon realized that his true research interests lay in Developmental and Evolutionary Biology. After graduating, Ben moved to Missoula, Montana, with nine of his friends, where he worked in Doug Emlen's lab studying the development and evolutionary diversification of horned beetles. Ben joined the Extavour lab in the Fall of 2008, and hopes to help figure out how the earliest stages of egg patterning vary between arthropod species possessing different ovary types.

Ben is a graduate student in the OEB program, and holds a National Science Foundation Graduate Research Fellowship. He successfully defended his Ph.D. thesis on April 18, 2014. Congratulations Ben! See photos of the champagne toasts here.


2014. Donoughe, S., Nakamura, T., Ewen-Campen, B., Green II, D.A., Henderson, L. and Extavour, C.G. BMP signalling is required for generation of primordial germ cells in an insect. Proceedings of the National Academy of Sciences of the USA 111(11): 4133-4138

2013. Germ cell specification requires zygotic mechanisms rather than germ plasm in a basally branching insect. Ewen-Campen, B., Donoughe, S., Clarke, D.N., and Extavour, C.G. Current Biology 23(10): 835-842

2013. Gene discovery in a hemimetabolous insect: a de novo transcriptome for the cricket Gryllus bimaculatus. Zeng, V. Ewen-Campen, B., Horch, H.W., Roth, S., Mito, T., and Extavour, C.G. PLoS ONE 8(5): e61479

2013. Evidence against a germ plasm in the milkweed bug Oncopeltus fasciatus, a hemimetabolous insect. Ewen-Campen, B.*, Jones, T.*, and Extavour, C.G. Biology Open (Company of Biologists) 2(6):556-568 (* equal author contribution)

2012. oskar Predates the Evolution of Germ Plasm in Insects. Ewen-Campen, B., Srouji, J.R., Schwager, E.E. and Extavour, C.G. Current Biology 22(23): 2278-2283. Read more in the Harvard Gazette and a Dispatch in Current Biology. Recommended by the Faculty of 1000.

2011. De novo assembly and characterization of a maternal and developmental transcriptome for the emerging model crustacean Parhyale hawaiensis. Zeng, V., Villanueva, K.E., Ewen-Campen, B., Alwes, F., Browne, W.E. and Extavour, C.G. BCM Genomics 12(1): 581.

2011. Delta/Notch signalling is not required for segment generation in the basally branching insect Gryllus bimaculatus. Kainz, F., Ewen-Campen, B., Akam, M. and Extavour, C.G. Development 138(22): 5015-5026.

2011. The maternal and early embryonic transcriptome of the milkweed bug Oncopeltus fasciatus. Ewen-Campen B., Shaner N., Panfilio K.A., Suzuki Y., Roth, S. and Extavour C.G. BMC Genomics. 12(1): 61. *Highly Accessed*

2010. Habitat type is related to nest mass and fledging success of Arctic Warblers. Hagelin J.C., Perry M.L., Ewen-Campen B., Sikes D.S. and Sharbaugh S. Wilson Journal of Ornithology 122(4):699-705.

2010. The molecular machinery of germ line specification. Ewen-Campen B.*, Schwager E.E.*, and Extavour C.G. Molecular Reproduction and Development 77(1): 3-18. (* equal author contribution)

2007. On the origin and evolutionary diversification of beetle horns. Emlen D.J., Corley Lavine L., and Ewen-Campen B. PNAS 104: suppl 1: 8661-8668.

2007. The aerodynamics of flying carpets. Gilbert SF, Chandra A, Cohen N, Ewen-Campen B, LaSalle K, Lewis K, Mizutani M, and Wu C. In: “The Panda’s Black Box.” Ed. Comfort NC. John Hopkins University Press.

tropharium tubulin adjusted